2Japan halted exports to other countries for fear that agar supported their development of biowarfare weapons. A few years before, Nazi Germany allegedly tested the efficacy of biowarfare attacks with another curious microbe, Serratia marcescens, dubbed “the miracle bacterium.” According to a much-talked about report by investigative journalist Henry Wickham Steed titled “Aerial Warfare: Secret German Plans” members of a secret Luft-Gas-Angriff (Air Gas Attack) Department spread the S. marcescens in the subterranean train networks of Paris and London and measured its reach armed with Petri dishes and agar plates.
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It is difficult to know exactly when the shift from gelatine to agar occurred. As often happens for scientific breakthroughs, agar was likely adopted incrementally alongside the use of other growth media. In 1913, for example, the first diagnosis of Serratia marcescens as a human pathogen was made by growing it on agar as well as on potatoes.
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